N. gonorrhoeae, Legionella pneumophila, enteropathogenic E. coli (EPEC), and Gram-positive
species. Type IV pili can present a 6 nm diameter and up to some µm in length. The T4P are
subdivided into three subtypes: T4aP, T4bP Tad pili, and T4cP designated recently based on
a sequence of amino acids of pilin [19]. The type IVa pili (T4aP) are long, straight, and
present polar bundles characterized by the existence of a PiT retraction ATPase and are
associated with enteric pathogens [20]. The type IVbP is present in between 180–238 amino
acids and has a signal peptide of 15–30 amino acids leader sequence. Many important
biological functions have been attributed to the T4P in Gram-negative bacteria such as
adherence to host cells, pathogenesis, formation of biofilms and microcolony, auto-
aggregation, and electric transfer as nanowires (Figure 11.2) [19,21]. They are considered
important virulence factors for many human and animal pathogens. In addition, the T4P
carries out another function, which is related to locomotion, termed twitching and gliding
motility in many bacterial species [22].
11.3 Microbial Nanowires and Bacterial Extracellular Electron
Transfer (EET)
In 2005, Reguera et al. [23] described the “microbial nanowire,” for the first time in mi
crobiology literature. The bacterial nanowires are considered an important component in
microbe-electrode and microbe-microbe electron exchange. The microbial nanowires rang
up tens of µm long and facilitated long-range EET (Figure 11.3). In the field of electro
microbiology, the nanowires enabled the bacteria that has electrically conductive pili, to
be considered as one of the most important discoveries. The microbial nanowires can be
defined as extracellular proteins that transfer an electron from cell to extracellular sub
stances under physiological conditions [24,25].
FIGURE 11.2
Schematic illustrations of the different functions of type IV pili. Reprinted with permission [ 19]. Copyright ©
2020 Published by Elsevier, under a Creative Commons Attribution 4.0 International License (CC BY-NC-ND).
Microbial Nanowires
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